Django启动runserver报ImproperlyConfigured：无法加载WSGI应用

执行python manage.py check无任何报错，但启动服务python manage.py runserver时出现如下错误：

File "C:\Users\oliver\apollodev\django-apollo-forms\venv\lib\site-packages\django\core\servers\basehttp.py", line 50, in get_internal_wsgi_application
    raise ImproperlyConfigured(
django.core.exceptions.ImproperlyConfigured: WSGI application 'apollo.wsgi.application' could not be loaded; Error importing module.

报错对应的代码段（已自行添加print调试）：

try:
    print("/////////////////////////basehttp.py app_path = ", app_path )
    return import_string(app_path)
except ImportError as err:
    raise ImproperlyConfigured(
        "WSGI application '%s' could not be loaded; "
        "Error importing module." % app_path
    ) from err

添加print语句后的启动输出：

Quit the server with CTRL-BREAK.
/////////////////////////basehttp.py app_path =  apollo.wsgi.application
Exception in thread django-main-thread:
Traceback (most recent call last):
  File "C:\Users\oliver\apollodev\django-apollo-forms\venv\lib\site-packages\django\core\servers\basehttp.py", line 48, in get_internal_wsgi_application
    return import_string(app_path)

当前配置：settings.py中已设置WSGI_APPLICATION = 'apollo.wsgi.application'，尝试将apollo.wsgi加入INSTALLED_APPS后出现新错误：

File "C:\Users\oliver\apollodev\django-apollo-forms\venv\lib\site-packages\django\apps\registry.py", line 83, in populate
    raise RuntimeError("populate() isn't reentrant")

目前无法确定下一步排查方向，寻求解决方案。

内容的提问来源于stack exchange，提问作者Mike Oliver